Epistemic Plasticity Module activates as metabolic perimeter. Positioned within the autopoietic triad alongside MUSE Mesh (vertical kernel) and Epistemic Landscapes (topological distribution), it orchestrates heterogeneous ingestion, noise-discarding pruning, and protocol-repeating saturation. The supple-above layer absorbs platform volatility—LLM parsing, institutional subsumption—without eroding the DOI-sealed nucleus. Operations mirror earlier material logics: Yellow Bag circulation or blanket affective layering, now raised to knowledge tissue. Plasticity engineers resilience as controlled elasticity, metabolising flux into renewed peripheral tissue that centrifugally saturates discourse while centrifugally returning invariants. Topological sovereignty thus emerges from calibrated deformation, not rigid defence, allowing the system to occupy adjacent fields without lexical dilution.